We are interested in characterizing when and how different brain systems for learning interact and whether this interaction is competitive or cooperative.We focus on two main brain systems for learning - one in the striatum and the other in the hippocampus. Traditionally, the striatum and hippocampus were thought to support independent and distinct learning systems. We have demonstrated that there is cross-talk between these systems during learning, raising questions about the nature of this interaction and its significance for learning and decision making. We are now investigating how this cross-talk is impacted by several key factors, including: motivation and feedback, social context of learning, aging, and genetic differences between learners.